:root{--ink:#132027;--muted:#53636b;--teal:#1e6b70;--line:rgba(255,255,255,.34);--paper:rgba(248,250,249,.93);--paper-dark:rgba(9,20,25,.80);--shadow:0 22px 70px rgba(0,0,0,.25);--heading:"Meiryo","メイリオ","Yu Gothic","游ゴシック",sans-serif}
*{box-sizing:border-box}html{scroll-behavior:smooth}body{margin:0;color:var(--ink);font-family:"Yu Gothic","游ゴシック","Hiragino Kaku Gothic ProN",sans-serif;line-height:1.8;background:#061116 url("assets/hero-bg.webp") center center/cover fixed no-repeat;overflow-x:hidden}body:before{content:"";position:fixed;inset:0;background:linear-gradient(180deg,rgba(3,10,14,.28),rgba(3,10,14,.58));z-index:-1}.shell{width:min(1180px,calc(100% - 48px));margin-inline:auto}.site-header{position:sticky;top:0;z-index:20;background:rgba(5,14,18,.78);backdrop-filter:blur(18px);border-bottom:1px solid rgba(255,255,255,.14)}.header-inner{min-height:78px;width:min(1280px,calc(100% - 40px));margin:auto;display:flex;align-items:center;gap:34px;color:white}.brand{display:flex;flex-direction:column;color:white;text-decoration:none;line-height:1.1;margin-right:auto}.brand strong{font:700 17px/1.2 var(--heading);letter-spacing:.08em}.brand span{font:600 12px/1.2 var(--heading);color:#a9d6d7;margin-top:6px}.site-nav{display:flex;gap:26px}.site-nav a,.lang-switch button{color:white;text-decoration:none;font:600 12px var(--heading);letter-spacing:.06em;background:none;border:0}.lang-switch{display:flex;gap:7px;align-items:center;font-size:11px}.lang-switch button{opacity:.56;padding:4px}.lang-switch button.active{opacity:1;border-bottom:1px solid #70c7ca}.menu-toggle{display:none;background:none;border:0;color:white;font-size:24px}.hero{min-height:750px;display:flex;align-items:center;padding:120px 0 95px}.hero-copy{width:min(1020px,100%);color:white;text-shadow:0 2px 18px rgba(0,0,0,.58)}.site-id,.kicker{font-family:var(--heading);font-weight:700;letter-spacing:.12em;color:#8ed4d6;margin:0 0 18px}.site-id{font-size:clamp(1.2rem,2vw,1.85rem)}.hero h1{font-family:var(--heading);font-weight:700;line-height:1.38;letter-spacing:.02em;margin:0 0 28px;font-size:clamp(2.5rem,5vw,4.9rem)}.hero-line{display:block;white-space:nowrap}.hero-lead{max-width:1000px;font-family:var(--heading);font-weight:500;font-size:clamp(1rem,1.55vw,1.3rem);color:rgba(255,255,255,.94);margin:0 0 34px}.affiliation-mark{display:flex;align-items:center;gap:22px;width:max-content;max-width:100%;padding:16px 20px;background:rgba(5,15,20,.58);border-left:3px solid #79c6c9;backdrop-filter:blur(8px)}.affiliation-mark img{width:180px;height:auto}.affiliation-mark strong,.affiliation-mark span{display:block;text-shadow:none}.affiliation-mark strong{font:700 16px var(--heading)}.affiliation-mark span{font:500 13px var(--heading);color:rgba(255,255,255,.84);margin-top:4px}.panel{border:1px solid var(--line);box-shadow:var(--shadow);backdrop-filter:blur(15px);background:var(--paper);padding:42px 48px;margin-bottom:28px}.news{margin-top:-48px;position:relative;z-index:3}.section-title-row{display:flex;justify-content:space-between;gap:30px;align-items:flex-end;margin-bottom:28px}.section-title-row h2,.profile h2{font-family:var(--heading);font-weight:700;font-size:clamp(1.5rem,2.2vw,2.35rem);line-height:1.5;margin:0;letter-spacing:.01em}.large-kicker{font-size:1rem}.text-link,.detail-link{color:var(--teal);text-decoration:none;font-weight:700;font-size:13px}.news-list{border-top:1px solid #ccd3d3}.news-list a{display:grid;grid-template-columns:100px 108px 1fr;gap:18px;align-items:start;padding:22px 0;border-bottom:1px solid #d7dddd;color:inherit;text-decoration:none}.news-list time{font-family:var(--heading);color:#4f6269}.tag{font:700 10px var(--heading);letter-spacing:.08em;color:#27676b}.news-list strong{font-family:var(--heading);font-weight:700}.news-list p{margin:6px 0 0;color:var(--muted);line-height:1.7}.profile{display:grid;grid-template-columns:1.18fr .82fr;gap:48px;align-items:center}.profile-image{height:430px;overflow:hidden;background:white}.profile-image img{width:100%;height:100%;object-fit:cover;object-position:center 36%;filter:brightness(1.18) contrast(1.06) saturate(.92)}.profile-copy h2 small{display:block;font-size:.48em;font-weight:600;color:#49616a;margin-top:6px}.profile-copy .role{font-weight:700;color:#31545b}.research-grid{display:grid;grid-template-columns:repeat(3,1fr);gap:18px}.research-card{background:rgba(255,255,255,.72);border:1px solid #d4dddd}.image-slot{height:220px;display:flex;align-items:end;padding:18px;background-position:center;background-size:cover;position:relative;overflow:hidden}.image-slot:before{content:"";position:absolute;inset:0;background:linear-gradient(0deg,rgba(6,18,22,.78),rgba(6,18,22,.02))}.image-slot span{position:relative;color:white;font-size:12px;line-height:1.6}.slot-1{background-image:radial-gradient(circle at 35% 40%,rgba(113,201,203,.58),transparent 22%),linear-gradient(135deg,#13333b,#051116)}.slot-2{background-image:radial-gradient(circle at 60% 35%,rgba(207,167,113,.50),transparent 26%),linear-gradient(135deg,#303837,#101818)}.slot-3{background-image:radial-gradient(circle at 40% 55%,rgba(126,190,159,.56),transparent 20%),linear-gradient(135deg,#12313b,#061116)}.card-content{padding:24px}.number{font:700 12px var(--heading);color:#3e878b}.card-content h3,.application-grid h3{font-family:var(--heading);font-size:1.16rem;line-height:1.65;margin:9px 0 10px}.card-content p{font-size:14px}.detail-link{display:inline-block;margin-top:8px}.section-note{font-size:12px;color:var(--muted)}.application-grid{display:grid;grid-template-columns:1fr 1fr;gap:22px}.application-grid article{background:rgba(255,255,255,.62);padding:18px}.image-slot.wide{height:260px}.methane{background-image:linear-gradient(0deg,rgba(3,20,29,.78),transparent),radial-gradient(circle at 70% 30%,#4d8498,transparent 25%),linear-gradient(145deg,#0d2935,#071218)}.mine{background-image:linear-gradient(0deg,rgba(20,26,22,.80),transparent),radial-gradient(circle at 35% 55%,#91855d,transparent 24%),linear-gradient(145deg,#38423a,#111715)}.publications .section-title-row h2{font-size:clamp(1.75rem,2.6vw,2.7rem)}.pub-list article{display:grid;grid-template-columns:90px 1fr;gap:24px;padding:22px 0;border-top:1px solid #ccd4d4}.pub-list article:last-child{border-bottom:1px solid #ccd4d4}.pub-list h3{font-size:15px;line-height:1.6;margin:0}.pub-list p{margin:5px 0 0;color:var(--muted)}.footer{display:grid;grid-template-columns:1.6fr .8fr .8fr;gap:36px;margin-top:28px;margin-bottom:40px;background:var(--paper-dark);color:white}.footer-brand{display:flex;gap:18px;align-items:flex-start}.footer-brand img{width:145px;height:auto}.footer span{display:block;color:#7fc1c4;font:700 10px var(--heading);letter-spacing:.12em;margin-bottom:8px}.footer a{display:block;color:white;text-decoration:none;margin:3px 0}.footer p{margin:4px 0 0;color:rgba(255,255,255,.78);font-size:13px}
@media(max-width:900px){.site-nav{display:none}.site-nav.open{display:flex;position:absolute;top:78px;left:0;right:0;padding:22px;flex-direction:column;background:#07151a}.menu-toggle{display:block}.hero{min-height:700px;padding-top:95px}.hero h1{font-size:clamp(2.3rem,7vw,4rem)}.hero-lead br{display:none}.affiliation-mark{width:100%}.profile,.research-grid,.application-grid{grid-template-columns:1fr}.profile-image{height:380px}.news-list a{grid-template-columns:80px 95px 1fr}.footer{grid-template-columns:1fr 1fr}.footer-brand{grid-column:1/-1}}
@media(max-width:620px){.shell{width:calc(100% - 24px)}.header-inner{width:calc(100% - 24px)}.brand span{display:none}.hero{min-width:0;min-height:660px}.hero h1{font-size:2rem;line-height:1.45}.hero-line{white-space:nowrap}.site-id{font-size:1rem}.hero-lead{font-size:.98rem}.affiliation-mark{align-items:flex-start;flex-direction:column;padding:14px 16px}.affiliation-mark img{width:150px}.panel{padding:30px 24px}.section-title-row{align-items:flex-start;flex-direction:column}.news-list a{grid-template-columns:72px 1fr;gap:8px 12px}.news-list .tag{grid-column:2}.news-list div{grid-column:1/3}.footer{grid-template-columns:1fr}}

/* v4 refinements */
.news .section-title-row { margin-bottom: 0.7rem; }
.news-link-cue {
  display: inline-flex;
  align-items: center;
  margin-top: 0.55rem;
  font-weight: 700;
  color: var(--accent, #176f78);
  border-bottom: 1px solid currentColor;
  padding-bottom: 0.08rem;
}
.news-list > a:hover .news-link-cue,
.news-list > a:focus-visible .news-link-cue { opacity: 0.72; }
.profile-copy .role { line-height: 1.85; margin-bottom: 1rem; }
.footer a[target="_blank"] { text-decoration: underline; text-underline-offset: 0.22em; }

/* v5: authentic microscopy image cards */
.research-image{aspect-ratio:16/8.2;overflow:hidden;background:#10191d;border-bottom:1px solid rgba(255,255,255,.12)}
.research-image img{width:100%;height:100%;display:block;object-fit:cover;transition:transform .6s ease,filter .6s ease;filter:saturate(.9) contrast(1.03)}
.research-card:hover .research-image img{transform:scale(1.025);filter:saturate(1) contrast(1.06)}

/* publication page */
.publication-page{padding-top:118px}
.publication-hero{padding:70px 54px 42px;color:#fff}
.publication-hero h1{font-family:Meiryo,"メイリオ",sans-serif;font-size:clamp(2.3rem,5vw,4.8rem);line-height:1.12;margin:.25em 0 .22em;letter-spacing:.02em}
.publication-hero>p:not(.kicker){font-size:clamp(.95rem,1.5vw,1.18rem);opacity:.85}
.publication-actions{display:flex;gap:24px;margin-top:26px;flex-wrap:wrap}
.publication-actions .text-link{color:#fff}
.publication-controls{display:flex;gap:8px;flex-wrap:wrap;padding:20px 24px;margin-bottom:18px}
.filter-button{border:1px solid rgba(10,32,38,.2);background:rgba(255,255,255,.55);border-radius:999px;padding:9px 16px;font:600 .79rem Meiryo,"メイリオ",sans-serif;cursor:pointer;transition:.2s}
.filter-button:hover,.filter-button.active{background:#163f45;color:#fff;border-color:#163f45}
.achievements{padding:42px 48px 58px}
.achievement-section+ .achievement-section{margin-top:64px;padding-top:52px;border-top:1px solid rgba(10,32,38,.16)}
.achievement-heading{display:grid;grid-template-columns:1fr auto;align-items:end;margin-bottom:22px}
.achievement-heading .kicker{grid-column:1/2;margin-bottom:7px;font-size:.95rem}
.achievement-heading h2{grid-column:1/2;font-family:Meiryo,"メイリオ",sans-serif;font-size:clamp(1.3rem,2.2vw,2rem);margin:0}
.achievement-heading>span{grid-column:2/3;grid-row:1/3;align-self:end;font-size:.78rem;opacity:.55}
.achievement-list{display:grid;gap:1px;background:rgba(10,32,38,.12);border-top:1px solid rgba(10,32,38,.12);border-bottom:1px solid rgba(10,32,38,.12)}
.achievement-item{display:grid;grid-template-columns:92px 1fr;gap:18px;background:rgba(250,251,249,.95);padding:18px 20px}
.achievement-item:hover{background:#fff}
.achievement-meta{display:flex;align-items:baseline;gap:10px;color:#326e73}
.achievement-number{font-weight:700;font-size:.8rem;opacity:.65}
.achievement-meta time{font-weight:700;font-size:1rem}
.achievement-item p{margin:0;line-height:1.72;font-size:.91rem}
.site-nav a.active{color:#75c4c7}
@media(max-width:720px){.publication-page{padding-top:82px}.publication-hero{padding:48px 22px 28px}.publication-controls{padding:16px}.achievements{padding:30px 16px 42px}.achievement-item{grid-template-columns:1fr;gap:8px;padding:16px}.achievement-meta{justify-content:space-between}.achievement-heading{display:block}.achievement-heading>span{display:block;margin-top:7px}.research-image{aspect-ratio:16/9}}

/* v6 additions */
.profile-statement {
  margin: 1.35rem 0 1.6rem;
  padding-top: 1.15rem;
  border-top: 1px solid rgba(20, 64, 75, 0.18);
  font-size: clamp(0.94rem, 1.35vw, 1.08rem);
  line-height: 1.95;
  color: #223b43;
}
.application-image {
  width: 100%;
  aspect-ratio: 16 / 8.2;
  overflow: hidden;
  border-radius: 12px;
  background: #071822;
  box-shadow: inset 0 0 0 1px rgba(92, 163, 170, 0.24);
  margin-bottom: 1.15rem;
}
.application-image img {
  display: block;
  width: 100%;
  height: 100%;
  object-fit: cover;
  transition: transform 0.65s ease;
}
.application-grid article:hover .application-image img { transform: scale(1.018); }
.publication-link {
  display: inline-block;
  margin-left: 0.45rem;
  font-weight: 700;
  color: #126a72;
  text-decoration: underline;
  text-underline-offset: 0.2em;
}

.doi-link{display:inline-block;margin:10px 0 0 74px;font-size:.84rem;font-weight:700;color:#176d73;text-decoration:underline;text-underline-offset:3px}.doi-search{color:#657579}.email-note{display:block;margin-top:4px;opacity:.72}.profile-page{padding-top:110px}.profile-hero,.profile-section{margin-bottom:18px;padding:clamp(24px,4vw,52px)}.profile-hero h1{font-family:Meiryo,"メイリオ",sans-serif;font-size:clamp(2rem,4vw,3.4rem);margin:8px 0 30px}.profile-hero h1 span{display:block;font-size:.42em;letter-spacing:.06em;margin-top:10px}.profile-detail-grid{display:grid;grid-template-columns:minmax(260px,.8fr) 1.2fr;gap:36px;align-items:center}.profile-photo-wrap img{width:100%;max-height:520px;object-fit:cover;border-radius:4px;filter:brightness(1.08) contrast(1.02)}.profile-facts dl{display:grid;grid-template-columns:6em 1fr;gap:18px 22px;margin:0}.profile-facts dt{font-weight:800;color:#146c72}.profile-facts dd{margin:0;line-height:1.9}.profile-section h2{font-family:Meiryo,"メイリオ",sans-serif;font-size:clamp(1.6rem,3vw,2.3rem);margin:7px 0 25px}.timeline,.award-list{list-style:none;margin:0;padding:0}.timeline li,.award-list li{display:grid;grid-template-columns:9em 1fr;gap:24px;padding:17px 0;border-top:1px solid rgba(5,35,45,.14)}.timeline time,.award-list time{font-weight:800;color:#146c72}@media(max-width:760px){.profile-detail-grid{grid-template-columns:1fr}.profile-facts dl,.timeline li,.award-list li{grid-template-columns:1fr}.doi-link{margin-left:0}}

/* v8: keep DOI labels on one line and align them with citation text */
.achievement-item .doi-link{grid-column:2;margin:8px 0 0;display:inline-block;width:max-content;max-width:100%;white-space:nowrap;overflow-wrap:normal;word-break:normal}
@media(max-width:720px){.achievement-item .doi-link{grid-column:1;margin-left:0;white-space:normal}}

/* v9: research page and year-grouped publications */
.profile-research-statement{margin-top:38px;padding:28px 30px;border-left:3px solid #5eabb0;background:rgba(15,66,73,.06)}
.profile-research-statement h2{margin:4px 0 12px;font:700 clamp(1.25rem,2.1vw,1.8rem)/1.55 var(--heading)}
.profile-research-statement p{margin:0;line-height:2;color:#29434b}
.research-page{padding-top:78px}
.research-hero{min-height:560px;display:flex;flex-direction:column;justify-content:center;color:#fff;text-shadow:0 2px 20px rgba(0,0,0,.55)}
.research-hero h1{font:700 clamp(2.7rem,6.5vw,5.8rem)/1.28 var(--heading);letter-spacing:.025em;margin:8px 0 22px}
.research-hero>p:last-child{font-size:clamp(1rem,1.45vw,1.25rem);line-height:2}
.research-section-head{max-width:900px;margin-bottom:34px}.research-section-head h2,.hunter-copy h2,.research-next h2{font:700 clamp(1.8rem,3.3vw,3rem)/1.45 var(--heading);margin:5px 0 18px}.section-intro{font-size:1.04rem;line-height:2;color:#334d55}
.cultivation-gap-grid{display:grid;grid-template-columns:.78fr 1fr 1.35fr;gap:40px;align-items:center}
.gtdb-stat{text-align:center;padding:28px;border:1px solid rgba(23,91,98,.18);background:rgba(255,255,255,.56)}.gtdb-stat .stat-label{display:block;font-size:.72rem;letter-spacing:.08em;color:#4f6f75}.gtdb-stat>strong{display:block;font:700 clamp(4.5rem,8vw,7rem)/1 var(--heading);color:#176b72;margin:12px 0 5px}.gtdb-stat>span:last-of-type{font-weight:700}.domain-counts{display:flex;justify-content:center;gap:34px;margin-top:24px;padding-top:18px;border-top:1px solid rgba(20,74,80,.15)}.domain-counts b,.domain-counts small{display:block}.domain-counts b{font-size:1.5rem}.domain-counts small{color:#60757a}
.donut-wrap{text-align:center}.donut{width:min(270px,100%);aspect-ratio:1;margin:auto;border-radius:50%;background:conic-gradient(#163e46 0 70%,#6db7ba 70% 100%);display:grid;place-items:center;position:relative;box-shadow:0 14px 40px rgba(0,0,0,.13)}.donut:before{content:"";position:absolute;width:64%;height:64%;border-radius:50%;background:#f5f7f5}.donut>div{position:relative;z-index:1}.donut strong,.donut span{display:block}.donut strong{font:700 2.6rem var(--heading);color:#173f46}.donut span{font-size:.8rem;color:#52666b}.donut-legend{display:flex;justify-content:center;gap:18px;flex-wrap:wrap;margin-top:18px;font-size:.78rem}.donut-legend span:before{content:"";display:inline-block;width:10px;height:10px;border-radius:50%;margin-right:7px}.legend-cultured:before{background:#6db7ba}.legend-uncultured:before{background:#163e46}
.gap-copy p{margin:0 0 16px;line-height:1.95}.source-link{color:#176d73;font-weight:700;text-decoration:underline;text-underline-offset:4px}.research-note{font-size:.74rem;color:#617378;margin:25px 0 0}
.unknown-grid{display:grid;grid-template-columns:repeat(3,1fr);gap:18px}.unknown-grid article{padding:28px;background:rgba(255,255,255,.6);border-top:3px solid #4f969b}.icon-mark{font:700 .8rem var(--heading);color:#398187}.unknown-grid h3{font:700 1.12rem/1.55 var(--heading);margin:10px 0}.unknown-grid p{font-size:.9rem;line-height:1.85;margin:0;color:#40575e}
.hunter-section{display:grid;grid-template-columns:1fr 1.2fr;gap:52px;padding:70px 48px;margin-bottom:28px;color:#fff;background:rgba(5,19,24,.84);border:1px solid rgba(255,255,255,.2);box-shadow:var(--shadow);backdrop-filter:blur(15px)}.hunter-copy p{line-height:1.95;color:rgba(255,255,255,.84)}.discovery-path{display:grid;grid-template-columns:1fr auto 1fr;gap:12px;align-items:center}.discovery-path div{min-height:130px;padding:20px;border:1px solid rgba(126,202,205,.35);background:rgba(255,255,255,.06)}.discovery-path i{font-style:normal;color:#75c4c7}.discovery-path span,.discovery-path strong,.discovery-path small{display:block}.discovery-path span{color:#75c4c7;font-weight:700}.discovery-path strong{font-family:var(--heading);margin:8px 0}.discovery-path small{color:rgba(255,255,255,.66)}
.next-grid{display:grid;grid-template-columns:repeat(3,1fr);gap:16px}.next-grid div{padding:22px;border:1px solid rgba(20,80,87,.16);background:rgba(255,255,255,.58)}.next-grid span,.next-grid strong,.next-grid em{display:block}.next-grid span{font-size:.75rem;color:#397e83}.next-grid strong{font-family:var(--heading);margin:8px 0}.next-grid em{font-size:.82rem;color:#5d7075}
.publication-controls{display:none}.achievement-section + .achievement-section{margin-top:48px;padding-top:42px}
@media(max-width:900px){.cultivation-gap-grid{grid-template-columns:1fr 1fr}.gap-copy{grid-column:1/-1}.unknown-grid{grid-template-columns:1fr}.hunter-section{grid-template-columns:1fr}.next-grid{grid-template-columns:1fr}.discovery-path{grid-template-columns:1fr 1fr}.discovery-path i{display:none}}
@media(max-width:620px){.research-hero{min-height:500px}.cultivation-gap-grid{grid-template-columns:1fr}.hunter-section{padding:42px 24px}.discovery-path{grid-template-columns:1fr}.profile-research-statement{padding:22px 18px}.profile-research-statement br{display:none}}

/* v10: Research page based on the supplied MOTIVATION layout */
.research-motivation-page{padding-top:92px;padding-bottom:48px}.motivation-panel,.featured-bacteria-section{background:rgba(246,247,247,.95)}.motivation-panel{padding:34px 42px 40px}.motivation-top{display:grid;grid-template-columns:minmax(0,1.55fr) minmax(280px,.75fr);gap:56px;align-items:center}.motivation-copy{font-size:1.02rem;line-height:1.9;color:#26363a}.motivation-copy p{margin:0 0 1.25em}.motivation-chart{position:relative;min-height:330px;display:grid;place-items:center}.donut-186{width:255px;height:255px;background:conic-gradient(#194f53 0 70%,#7fc7c4 70% 100%);border-radius:50%;display:grid;place-items:center;position:relative}.donut-186:before{content:"";position:absolute;width:128px;height:128px;border-radius:50%;background:#f6f7f7}.donut-186>div{position:relative;z-index:1;text-align:center;display:flex;flex-direction:column}.donut-186 strong{font-size:3rem;line-height:1;color:#155f62}.donut-186 span{font-size:.72rem;margin-top:.45rem;color:#53676a}.donut-side-label{position:absolute;font-size:.92rem;line-height:1.45;color:#34484b}.donut-side-label b{font-size:1.05rem;color:#155f62}.cultured-label{left:-6px;top:48px;text-align:right}.uncultured-label{right:0;bottom:34px}.featured-bacteria-section{padding:30px 42px 54px;margin-top:22px}.featured-bacterium{display:grid;grid-template-columns:minmax(0,1fr) minmax(360px,.92fr);gap:48px;align-items:center;padding:36px 0;border-top:1px solid rgba(31,67,69,.16)}.featured-bacterium:first-of-type{margin-top:10px}.featured-bacterium-copy h2{font-family:Meiryo,"メイリオ",sans-serif;font-size:clamp(1.15rem,1.55vw,1.55rem);line-height:1.55;margin:0 0 8px;color:#26383a}.featured-bacterium-copy .species{margin:0 0 16px;color:#397579}.featured-bacterium-copy>p:not(.species){font-size:.96rem;line-height:1.9;color:#45595c;margin:0 0 18px}.featured-bacterium-image{display:block;background:#fff;padding:6px;border:1px solid rgba(25,80,84,.18);box-shadow:0 8px 28px rgba(7,30,37,.08);overflow:hidden}.featured-bacterium-image img{display:block;width:100%;height:auto;max-height:330px;object-fit:contain;background:#fff;transition:transform .25s ease}.featured-bacterium-image:hover img{transform:scale(1.015)}.research-links{display:flex;flex-wrap:wrap;gap:10px 18px}.research-links a{color:#126b70;text-decoration:none;border-bottom:1px solid currentColor;font-weight:700;font-size:.86rem}.research-links a:hover{opacity:.72}.publication-all .achievement-heading h2{font-size:clamp(2rem,4vw,3.4rem)}
@media(max-width:800px){.motivation-top,.featured-bacterium{grid-template-columns:1fr}.motivation-top{gap:18px}.motivation-panel,.featured-bacteria-section{padding-left:24px;padding-right:24px}.motivation-chart{min-height:280px}.featured-bacterium{gap:22px}.featured-bacterium-image{order:-1}.cultured-label{left:4px}.uncultured-label{right:4px}}

/* v11: publication years without sequential numbering; richer Energy & Environment summaries */
.publication-all .achievement-meta { min-width: 72px; }
.publication-all .achievement-meta time { font-weight: 700; color: var(--teal, #287f84); font-size: 1.05rem; }
.application-grid article > p { line-height: 1.9; }
.application-grid .detail-link[target="_blank"]::after { content: ""; }


/* Public release polish */
:root{--focus:#91d9db;--radius:4px;--content-max:1180px}
html{font-size:16px;text-rendering:optimizeLegibility;-webkit-font-smoothing:antialiased}
body{background-image:url("assets/hero-bg.webp");}
a{transition:color .18s ease,opacity .18s ease,border-color .18s ease,background-color .18s ease}
a:focus-visible,button:focus-visible{outline:3px solid var(--focus);outline-offset:4px;border-radius:2px}
.site-nav a{position:relative;padding:7px 0}.site-nav a:after{content:"";position:absolute;left:0;right:100%;bottom:0;height:1px;background:#76c6c9;transition:right .2s ease}.site-nav a:hover:after,.site-nav a.active:after{right:0}
.panel{border-radius:var(--radius)}
.section-title-row h2,.profile h2,.profile-section h2,.featured-bacterium-copy h2{letter-spacing:.005em}
.hero{min-height:min(760px,92vh)}
.hero h1{font-size:clamp(2.35rem,4.5vw,4.55rem);max-width:1100px}
.news-list a:hover{background:rgba(63,138,143,.045)}
.research-card,.application-grid article{transition:transform .22s ease,box-shadow .22s ease,border-color .22s ease}
.research-card:hover,.application-grid article:hover{transform:translateY(-2px);box-shadow:0 14px 34px rgba(10,38,45,.09);border-color:rgba(34,111,117,.34)}
.research-image img,.application-image img{display:block;width:100%;height:100%;object-fit:cover}
.text-link,.detail-link,.research-links a,.source-link,.doi-link{text-decoration-thickness:1px;text-underline-offset:4px}
.footer{margin-top:42px}
.reveal{opacity:0;transform:translateY(12px);transition:opacity .48s ease,transform .48s ease}.reveal.is-visible{opacity:1;transform:none}

/* Refined GTDB composition graphic */
.motivation-chart{isolation:isolate}.donut-186{width:270px;height:270px;background:conic-gradient(#194f53 0 69.5%,#83cac8 69.5% 100%);box-shadow:0 16px 46px rgba(12,54,61,.16),inset 0 0 0 1px rgba(255,255,255,.55)}
.donut-186:after{content:"~70% uncultivated";position:absolute;right:-7px;bottom:-33px;font:700 .72rem var(--heading);letter-spacing:.03em;color:#194f53;background:rgba(246,247,247,.92);padding:4px 7px;border-radius:2px}
.donut-186>div{background:#f7f8f8;width:134px;height:134px;border-radius:50%;display:grid;place-content:center;box-shadow:0 2px 12px rgba(0,0,0,.05)}
.donut-186:before{display:none}.donut-186 strong{font-size:3.15rem}.donut-186 span{max-width:95px;line-height:1.35;margin-inline:auto}.donut-side-label{background:rgba(247,248,248,.88);padding:6px 8px;border-radius:2px}.cultured-label{left:0;top:52px}.uncultured-label{right:-2px;bottom:36px}
html[lang="en"] .donut-186:after{content:"~70% uncultivated"}
html[lang="ja"] .donut-186:after{content:"約70% 未培養"}

/* Publications: calmer continuous bibliography */
.publication-all .achievement-list{border-top:1px solid rgba(14,56,64,.18)}
.publication-all .achievement-item{grid-template-columns:72px minmax(0,1fr);column-gap:30px;padding:24px 8px;border-bottom:1px solid rgba(14,56,64,.13)}
.publication-all .achievement-item p{margin:0;line-height:1.8}
.publication-all .achievement-item .doi-link{grid-column:2;margin-top:7px;font-size:.9rem;width:auto;max-width:100%;white-space:normal;overflow-wrap:anywhere}
.publication-all .achievement-meta{padding-top:1px}
.publication-all .achievement-meta time{font-size:1rem}
.shared-publications-note{max-width:680px;color:var(--muted)}

/* Error page */
.error-page{min-height:100vh;display:grid;place-items:center;padding-block:90px}.error-card{width:min(760px,100%);text-align:center;margin:0}.error-card h1{font:700 clamp(2rem,5vw,4rem)/1.2 var(--heading);margin:.2em 0}.error-card p{color:var(--muted)}.error-actions{display:flex;gap:28px;justify-content:center;flex-wrap:wrap;margin-top:28px}

@media(max-width:900px){.hero{min-height:680px}.site-nav{gap:18px}.header-inner{gap:22px}.motivation-top{gap:28px}}
@media(max-width:760px){
  .shell{width:min(100% - 28px,1180px)}.header-inner{width:min(100% - 28px,1280px);min-height:68px}.menu-toggle{display:block;margin-left:auto}.lang-switch{order:3}.site-nav{display:none;position:absolute;top:68px;left:0;right:0;background:rgba(5,14,18,.97);padding:16px 24px 22px;border-top:1px solid rgba(255,255,255,.1);flex-direction:column;gap:3px}.site-nav.open{display:flex}.site-nav a{padding:12px 0}.brand{margin-right:0}.brand strong{font-size:15px}
  .hero{min-height:610px;padding:90px 0 68px}.hero h1{font-size:clamp(2rem,9vw,3rem)}.hero-line{white-space:normal}.affiliation-mark{width:100%;padding:14px 14px;gap:14px}.affiliation-mark img{width:120px}.affiliation-mark strong{font-size:13px}.affiliation-mark span{font-size:11px}
  .panel{padding:30px 24px}.news{margin-top:-24px}.news-list a{grid-template-columns:80px 1fr}.news-list .tag{grid-column:1}.news-list a>div{grid-column:2;grid-row:1/3}
  .profile{grid-template-columns:1fr;gap:24px}.profile-image{height:380px}.research-grid,.application-grid{grid-template-columns:1fr}.research-card:hover,.application-grid article:hover{transform:none}
  .motivation-panel,.featured-bacteria-section{padding-left:22px;padding-right:22px}.motivation-chart{min-height:310px}.donut-186{width:235px;height:235px}.donut-186>div{width:118px;height:118px}.donut-186 strong{font-size:2.7rem}.donut-side-label{font-size:.8rem}.cultured-label{left:-2px}.uncultured-label{right:-2px}
  .publication-all .achievement-item{grid-template-columns:1fr;padding:22px 0}.publication-all .achievement-item .doi-link{grid-column:1}.publication-all .achievement-meta{margin-bottom:6px}
}
@media(prefers-reduced-motion:reduce){html{scroll-behavior:auto}*,*:before,*:after{animation-duration:.001ms!important;transition-duration:.001ms!important;scroll-behavior:auto!important}.reveal{opacity:1;transform:none}}

/* v1.1 readability and layout refinements */
.header-inner{min-height:88px;gap:40px}
.brand strong{font-size:26px;letter-spacing:.055em}
.brand span{font-size:14px;margin-top:7px;letter-spacing:.055em;color:#bde8e8}
.site-nav{gap:30px}
.site-nav a{font-size:16px;letter-spacing:.045em;padding:10px 0;color:#f6fbfb}
.lang-switch{gap:9px;font-size:16px;font-family:var(--heading);font-weight:800}
.lang-switch a{color:#d8f4f4;text-decoration:none;padding:8px 7px;border-radius:4px;line-height:1}
.lang-switch a:hover{color:#fff;background:rgba(126,213,216,.14)}
.lang-switch a.active{color:#7fe0e3;border-bottom:2px solid #7fe0e3}
.lang-switch span{color:rgba(255,255,255,.55)}

/* Let the hero artwork breathe: content stays in the upper-left and no opaque hero box. */
.hero{min-height:min(800px,92vh);align-items:flex-start;padding:92px 0 150px}
.hero-copy{width:min(820px,100%)}
.site-id{font-size:clamp(1.8rem,2.8vw,2.65rem);margin-bottom:22px;color:#a5e1e3;letter-spacing:.09em}
.hero h1{font-size:clamp(2.35rem,4.25vw,4.25rem);margin-bottom:34px}
.affiliation-mark{background:transparent;border-left:0;backdrop-filter:none;padding:0;gap:20px}
.affiliation-mark img{width:155px}
.affiliation-mark strong{font-size:15px;text-shadow:0 2px 12px rgba(0,0,0,.75)}
.affiliation-mark span{font-size:12.5px;color:rgba(255,255,255,.92);text-shadow:0 2px 12px rgba(0,0,0,.75)}

/* Stronger section cues on the home page. */
.large-kicker{font-size:1.35rem;letter-spacing:.11em;color:#176f74;margin-bottom:12px}
.news .large-kicker,.research .large-kicker{font-size:1.55rem;color:#0c646a}

/* Rebalance home Profile: smaller image, more room for identity and affiliation. */
.profile{grid-template-columns:minmax(250px,.72fr) minmax(0,1.28fr);gap:52px}
.profile-image{height:315px;max-width:430px;justify-self:center;width:100%}
.profile-copy{align-self:center}
.profile-copy h2{font-size:clamp(1.75rem,2.5vw,2.55rem)}

/* Contact information */
.contact-address{margin:12px 0 0;color:rgba(255,255,255,.82);font-size:12.5px;line-height:1.7;max-width:260px}

@media(max-width:900px){
  .header-inner{min-height:76px;gap:24px}
  .brand strong{font-size:21px}.brand span{font-size:12px}
  .lang-switch{font-size:15px}
  .hero{padding:80px 0 105px;min-height:680px}
  .profile{grid-template-columns:1fr}.profile-image{height:340px;max-width:500px}
}
@media(max-width:620px){
  .header-inner{min-height:70px}
  .brand strong{font-size:17px}.brand span{display:block;font-size:10px;margin-top:4px}
  .lang-switch{font-size:14px;gap:3px}.lang-switch a{padding:7px 5px}
  .hero{padding:72px 0 90px;min-height:610px}.site-id{font-size:1.35rem}.hero h1{font-size:2rem}
  .affiliation-mark img{width:130px}.affiliation-mark{gap:14px}
  .large-kicker,.news .large-kicker,.research .large-kicker{font-size:1.25rem}
  .profile-image{height:300px}
}